After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galápagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died. Gramma was born ...

Born in the 1880s, Gramma the Galapagos tortoise lived through world wars and pandemics while becoming a symbol of ...

LOS ANGELES -- After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died.

LOS ANGELES — After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died.

LOS ANGELES (AP) — After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galápagos tortoise Gramma, the oldest resident of the San Diego Zoo, has ...

Add Yahoo as a preferred source to see more of our stories on Google. LOS ANGELES (AP) — After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved ...

LOS ANGELES(AP) — After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died ...

LOS ANGELES (AP) — After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise Gramma, the oldest resident of the San Diego Zoo, has ...

No one knows exactly when Gramma the Galápagos tortoise was born on that volcanic chain of Pacific islands. What is clear, though, is that she lived through the fall of empires, two world wars, and ...