(FOX 5/KUSI) — Gramma, a Galapagos tortoise that was the oldest living animal at the San Diego Zoo, has died. The zoo said Gramma had ongoing bone conditions and that caretakers had to make the ...

SAN DIEGO (KGTV) — The San Diego Zoo announced on Friday via an Instagram post that Gramma, the beloved Galápagos tortoise, has died. Gramma arrived at the San Diego Zoo between 1928 and 1931 as part ...

San Diego Zoo staff and animal lovers are reeling this week following the death of beloved Gramma the Galápagos tortoise, by far the zoo's oldest resident, in what has been a year marked by tragic ...

Gramma, an estimated 141-year-old Galápagos tortoise, died Thursday at the San Diego Zoo, ending a near century-long chapter of what the zoo said was quiet, yet constant ambassadorship to generations ...