The San Diego Zoo's oldest resident, a Galapagos tortoise named Gramma, died Nov. 20. She was estimated to be 141 years old.

Gramma, the beloved Galápagos tortoise and the San Diego Zoo’s oldest resident, has died at an estimated age of 141. Known for her sweet, shy demeanour and a diet of romaine lettuce and cactus fruit, ...

San Diego Zoo mourns the loss of Gramma, a Galápagos tortoise estimated to be 141 years old. Born in the Galápagos Islands, ...

No one knows exactly when Gramma was born. But if the estimated birth year of 1884 is accurate, Chester Arthur occupied the Oval Office and there were only 39 states at the time.

Gramma, the beloved Galápagos tortoise and the San Diego Zoo’s oldest resident, has died at an estimated age of 141. Known for her sweet, shy demeanour and a diet of romaine lettuce and cactus fruit, ...

After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise ...

LOS ANGELES (AP) — After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise Gramma, the oldest resident of the San Diego Zoo, has ...

(WPHL) — For the first time in over 150 years, the Philadelphia Zoo has new additions to the Western Santa Cruz Galapagos tortoise family. The zoo announced the hatching of four critically endangered ...

Gramma the Galapagos tortoise was a roughly 141-year-old icon for visitors to the San Diego Zoo. After nearly a century of delighting guests, the zoo shared on social media this week that she died ...

After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died.Gramma was born ...

LOS ANGELES -- After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died.