Beloved Galápagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died at about 141, the zoo says, adding that "caring for such a remarkable tortoise was a privilege." ...

After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise ...

No one knows exactly when Gramma was born. But if the estimated birth year of 1884 is accurate, Chester Arthur occupied the Oval Office and there were only 39 states at the time.

San Diego Zoo mourns the loss of Gramma, a Galápagos tortoise estimated to be 141 years old. Born in the Galápagos Islands, ...

A 141-year-old tortoise, once celebrated as a star attraction at the San Diego Zoo in the United States, has passed away. According to multiple foreign media outlets including *The New York Times* on ...

The beloved Galapagos tortoise named Gramma that was the oldest resident of the San Diego Zoo at an estimated 141 years has died LOS ANGELES -- After more than a century of munching on her favorite ...

One Galápagos tortoise lived 141 years, turning quiet meals and slow steps into a legacy of care, memory, and conservation ...

The San Diego Zoo's oldest resident, a Galapagos tortoise named Gramma, died Nov. 20. She was estimated to be 141 years old.