The San Diego Zoo's oldest resident, a Galapagos tortoise named Gramma, died Nov. 20. She was estimated to be 141 years old.

The San Diego Zoo has said goodbye to one of its most cherished icons, Gramma, a Galápagos tortoise who inspired generations of visitors and conservationists. She passed away on November 20 after care ...

After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died. Gramma was born ...

San Diego Zoo mourns the loss of Gramma, a Galápagos tortoise estimated to be 141 years old. Born in the Galápagos Islands, she became a beloved \"Que ...

No one knows exactly when Gramma was born. But if the estimated birth year of 1884 is accurate, Chester Arthur occupied the ...

LOS ANGELES (AP) — After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise Gramma, the oldest resident of the San Diego Zoo, has ...