Conservationists are hoping to strengthen regulations at a wildlife trade conference on birds, reptiles and other animals ...

Gramma, who died at the zoo on Thursday, was believed to have been born in 1884, when Chester A. Arthur was in the White ...

If walls could talk, Jonathan the tortoise would have the longest story to tell because he has lived through centuries of ...

Her life spanned a tumultuous period of U.S. and world history, from Chester Arthur to Donald Trump. To San Diegans, she was ...

No one knows exactly when Gramma was born. But if the estimated birth year of 1884 is accurate, Chester Arthur occupied the ...

An occasional series where Mongabay founder Rhett Ayers Butler shares analysis, perspectives and story summaries. She moved ...

Gramma, an estimated 141-year-old Galápagos tortoise, died Thursday at the San Diego Zoo, ending a near century-long chapter ...

The San Diego Zoo's oldest resident, a Galapagos tortoise named Gramma, died Nov. 20. She was estimated to be 141 years old.

Gramma was described as "a quiet and constant presence at the zoo, a witness to history, a beloved icon, and an extraordinary ...

After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galápagos tortoise ...